Classificatioin and Phylogeny of the Animal Kingdom

- There are about 1.5 Million Species

Carolus Linnaeus:
Binomial nomenclature
Based on comparative anatomy
Kindgom, phylum, class, order, family, genus and species

How to Write a Scientific Name

1. First word is genus, written with capital initial letter
2. Second word is species, all small initial letter.
3. Both words are italic or underlines,
Eg:
Homo sapiens or Homo sapiens
Gorilla gorilla or Gorilla gorilla

Trypanosoma burcei rhodensiense
Trypanosoma brucei gambiense
Trypanosoma brucei brucei
in a sentence you might see this:
Trypanaosoma spp.
Gorilla gorilla or G. gorilla or G. gorilla
Trichomonas vaginalis or T. vaginalis or T. vaginalis

- Phylogeny is based on homology, not on analogy
- human
Rober Whitteker

Reproduction Main Events

Gamete Formation:
Fertilization:
Cleavage:
Gastrulation
Neurulation
Organogenesis
Growth

Embryology

- Fertilization
- Zygote - Parthenogenesie
Eg: rotifers, crustaceans, insects, fishes, desert lizards, etc
- Polyspermy

There are 4 Main Types of Cleavage:

1. Radial Holoblastic:

2. Spiral Holoblastic

3. Discoidal Meroblastic

4. Rotational Holoblastic

There is also SUPERFICIAL in insects

Cleavage in Invertebrates:

1. Radial or regulative
2. Spiral

- Protostomia: 'mouth-first'
- Deuterostomia: 'mouth-second'

There are 4 Types of Yolk:

1. Isolecithal: starfish and human
2. Mesolecithal: frog
3. Telolecithal: birds
4. Centrolecithal: insects

What is Cleavage?

: process of nucleus and cell division in zygote.
Blastula: is a hollow mass of cells
- Blastocoel

What is Gastrula?

: having 2 layers of cells with an opening
- Invagination:
- Archenteron:
- Coelomic Vesicles
- Now gastrula is an embryo and has 3 germ layers:
1. Ectoderm: body surface, epithelial cells, nervous system
2. Endodermis: epithelial cells of G.I. tract
3. Mesodermis: muscles, reproductive, peritoneum
-Diploblastic
- Triploblastic

Know the fate of blastopore in Protostomes and Deuterostomes

Formation of the Coelom

1. Schizocoelous
2. Enterocoelous
Advance chordates from Schizocoelous

Amniotes and the Amniotic Egg:

- Reptiles, birds, and mammals
- Amnion:
- Allantois:
- Chorion:
- Chorioallantoic membrane

33 Hours
Procen
Mesen
Rhomben
48 Hours
Prosen
Mesence
Myelence
72 Hours
Telence
Dience
Mesence
Metence
Myelence
96 Hours
Telence
Dience
Mesence
Metence
Myelen

Forebrains = prosencephalon
Midbrains = mesencephalon = include optic lobes
Hindbrain = rhombencephalon = medula

- Before 48 hours, there is porcen and NO dience
- After 48 hours there is prosen and there is deince
- At 72 hours procen becomes telence
- At 48 hours there is No telence

Animal Like Protista

- First life on earth about 3.5 billion years ago
- Protista kingdom
- Protozoa Subkingdon

Animal Like Because

A. No cell wall
B. In some way motile
C. Ingest their food

We will discuss 3 Phyla out of 7 Phuyla

Sarcomastigophora
Apicomplexa
Ciliophora

Main Characteristics of Protozoan Phyla:

1. Unicellular
2. Microscopic
3. No germ layers
4. Specialized organs
5. Free-living
Mutualism
Commensalism
Parasitism
6. Some sessile, pseudopodia, flagella, cilia
7. All types of nutrition: autotrophic, heterotrophic
8. Aquatic or terrestrial
9. Asexual and sexual reproduction

There have most of the organelles that other eucaryotic cells posses

- Mitochondria
- Endoplasmic Reticulum
- Golgi apparatus
- Vesicles
- Chloroplast
- Vacuoles
- Ectoplasm and endoplasm

Locomotion:

1. Cilia and flagella: 9+2
2. Pseudopodia:
Ectoplasm: transparent or hyaline, colliod
Endoplasm: more granular, more fluid
A. Lobopodia: large extension of cells body
B. Limax: whole body moves
C. Filopodia: thin extensions
D. Reticulopodia: rejoin to form a netlike mesh
E. Axopodia: supported by axial rods

Excretion and Osmoregulation:

By contractile vacuole: 1. Small vesicles join the contractile vacuole
2. Then contractile vacuole joins the cell membrane
In ciliates it is more complex:
1. Excretory pore leads to outside and they are surrounded by ampulla
2. Ampulla fills up and discharge the water to the outside

Nutrition:

A. Autotrophs: order Euglenida
B. Heterotrophs:
- Phagocytosis
- Lysosomes
- Food vacuole
- Exocytosis the waste material
- Cilia have cytoproct for expulsion of waste
- Pinocytosis

Reproduction:

A. Fission:
a. Binary fission:
b. Budding in cilia
c. Multiple fission
Sporogony: development of spores or sporozoites within occyst
Schizogony: when cytokinesis is preceded by several nuclear division

B. Sexual Reporduction
a. Isogametes
b. Anisogamete
c. Syngamy: fertilization of an individual gamete by another
d. Autogamy: gametic nuclei fuse to form zygote within the same organism that produced them
e. Conjugation: exchange of gametic nuclei between paired organisms

Encystment and Excystment:

Formation of cyst. Not in paramecium
Excape from cyst in called excystment

Phylum Sarcomastigophora:

Some mastigophorans can use psuedopodia and flagella.
Subphylum Mastigophora
A. Phytoflagellates
Eg: Euglena, Chlamydomonas and Volvox
Coral reefs are by zooxanthellae

Some Structures in Euglena:
- Pellicle
- Kinetosome
- Contractile vacuole
- Stigma to locate light?
- Chloroplast
- Paramylon bodies
- Reproduction by binary fission

Some Structures in Volvox:
- Cell wall
- Division of labor
- Asexual and sexual reproduction
- Macrogametes loaded with food
- Microgametes forms sperm to find ovum
- Zygoes multiply and break away

B. Zooflagellates:
Trypanosoma brucei rhodensiense
Trypanosoma brucei gambiese
Trypanosoma burcei brucei
transmitted by tsese fly (Glossina)
Trypanosoma cruzi cause Chagas' disease
- transmitted by kissing bug Triatominae
Leishmania in liver and spleen
- Visceral Leishmaniasis
- Cutaneous Leishmaniasis
- Transmitted by sand fly
Trichonomas vaginalis
Giardia lamblia

Subphylum Sarcodina

Entamoeba histolytica
Entamoeba coli
Entamoeba gingivalis
Forminiferas: shelled amoeba, make up of calcium carbonate, silica and silt radiolarians and heliozoan, silicon

Phylum Apicomplexa:

- Micronemes
- Pseudopodia in some intracellular stages
- Flagella in gametes of some species
- Asexual and sexual cycle
- Some require intermediate host
- Some develop oocyst or spore

Class Sporozoa:

- Most important class
Three Subclasses:
Gregarinia: found in insects
Coccidia:
Piroplasma: in cattle, Texas red-water fever caused by Babesia bigemina

Subclass Coccidia:

- Intracellular
- Eimeria and Isospora cause coccidiosis
- Eimeria tenella in young fowl
Reproduction of Eimeria:
Schizogony --> gametes --> fertilization --> oocyst --> in intestine in feces

Sporogony in oocyst --> 8 sporozoites --> outside of host new host ingest sporulated oocyst

Toxoplasma gondii
In AIDS patients
Definitive host is cat
Possible intermediate host

Plasmodium
Malaria transmitted by mosquitoes
Sporozoites injected to human by insects
Sporozoites in live initiate schizogony
P. falciparum penetrate R.B.C after one generation in liver
In RBC trophozoites feed on HBG
Sexual Reproduction
Microgamests and macrogametes
Mosquitoes ingest gametes
Fertilization --> oocyst

Phylum Ciliophora:

-Mostly freeliving some parasitic, some commensalism, and some colonial
- Some sessile; suctorians
- Larger than most protozoan
- Multinucleated
- Undulating membrane: cilia are fused into a sheet
- Cytostomes (mouth)
- Oral groove
- Two contractile vacuole system
Passage of water and food in paramecium:
Oral groove --> cytostome --> cytopharynx --> food vacuole --> indigestivle out by cytoproct

Reproduction

1. Binary fission: micronucleus divide mitotically and macronucleus elongate and divide amitotically
2. Conjugation
3. Autogamy: self-fertilization
eg: Balantidium coli

Phylogeny:

Apicomplexa from flagellated
Mastigophora from bacteria
Ciliated from flagellated or ?